The relationship between Black Greek-letter organization membership and African American student engagement in almost exclusively White college classrooms was explored in this study. Data were collected through interviews with 131 members from seven undergraduate chapters at a large, predominantly White university in the Midwest. This study resulted in an explanatory model that shows how underrepresentation, voluntary race representation, and collective responsibility positively affect active participation, while Forced Representation has a negative effect. Findings also reveal that faculty teaching styles both positively and negatively affect engagement among African American sorority and fraternity members in their classes. The implications of these findings are discussed at the end of the article. (Contains 1 figure.)