The current authors reply to a response by Bowers on a comment by the current authors on the original article. Bowers (2010) mischaracterizes the goals of parallel distributed processing (PDP research)--explaining performance on cognitive tasks is the primary motivation. More important, his claim that localist models, such as the interactive activation model, "recognize" their inputs when a threshold is reached runs directly counter to an essential feature of these models. This undermines his attempt to distinguish between what a neuron responds to and what it codes for and, indeed, undermines the whole localist argument he has proposed. Bowers's (2010) also continues to face difficulty in specifying what localist units correspond to, and all of the possible choices face problems. In the paragraphs below we substantiate these points.