This study presents a reanalysis of existing data to investigate whether a relationship between perception and production abilities regarding a challenging second-language (L2) phonological contrast is observable (a) when both modalities must rely on accessing stored lexical representations and (b) when there is an asymmetry in task focus between perception and production. In the original studies, German learners of English were tested on their mastery of the English /[open-mid front unrounded vowel]/-/ae/ contrast in an auditory lexical decision task with phonological substitutions, a word-reading task, and a segmentally focused imitation task. Results showed that accurate nonword rejection in the lexical decision task was predicted by the Euclidean distance between the two vowels in word reading but not in imitation. These results extend previous findings to lexical perception and production, highlight the influence of task focus on the degree of coupling between the two modalities, and may have important implications for pronunciation training methods.