In Carolyn Miller's "Rhetorical Community: The Cultural Basis of Genre," she revisits her assertion that genres are cultural artifacts and questions the nature of the relationship between micro-level, individual speech acts, and macro-level genres and systems. To demonstrate this relationship, I analyze "meta-genre" accounts of the dissertation posted on the Writing Program Administrator (WPA) listserv, a forum for Computer Mediated Communication (CMC). Within this discourse, I identify "transitional threads"--moments when the discussion shifts, which show the relationship between micro- and macro-level interaction on the listserv as well as constructions of the dissertation within Writing Studies. CMC highlights how micro-level speech acts aggregate and are impacted by macro-level culture, and it showcases the heterogeneity inherent in the rhetorical community of the listserv.